My favourite paragraph from The Human Zoo:

Physiological Sex can also be observed in other animal species and it is worth taking a look at a few examples. As one might expect, they are most readily encountered in the animal zoo, rather than in the wild state. Many zoo animals have been seen to masturbate when kept in isolation. This is most commonly observed in captive monkeys and apes. In males, the penis is stimulated sometimes by the hand or foot, sometimes by the mouth, and sometimes by the tip of the prehensile tail. Male elephants stimulate their penises with their trunks and female elephants kept in a group without a male stimulate one another’s genitals with their trunks. Even a male lion kept in isolation in a zoo cage has been seen to heave itself up into an inverted position against a wall and masturbate with its paws. Male porcupines have been observed to walk around on three legs, holding one forepaw on their genitals. One male dolphin developed the pattern of holding its erect penis in the powerful jet water intake of its pool. Sex dreaming also seems to occur in animals and in domestic cats, the erection of the penis while asleep has been observed to lead to full ejaculation.1

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1 Desmond Morris, The Human Zoo, 1969, p89

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